Uncharted 4: A Thief's End developer Naughty Dog has discussed the improvements made to its facial animation systems since Uncharted 3 and The Last of Us, saying that face models now animate with "between 300 and 500 bones."
In an interview with GamesTM , writer Josh Scherr contextualised the improvements by comparing them to previous Uncharted games and The Last of Us, which he said used "about 90 and 100 'bones' in their faces."

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